Chromosomes carry the Genes that Mendel studied. Cytology and the chromosomal theory of inheritance
Mendel's 2nd law vs chromosome theory How to resolve it: Fruit flies and linkage (experiments by Thomas Hunt Morgan, with Drosophila melanogaster
–> Genetic dominance (gives rise to modes of inheritance)
https://www.wikiwand.com/en/Boveri%E2%80%93Sutton_chromosome_theory